Cyanus triumfettii subsp. axillaris (Čelak.) Štěpánek (syn. Centaurea triumfetti subsp. aligera Dostál)

Taxonomy: Fam. Compositae// Genus:  Cyanus// Species: Cyanus triumfettii (All.) Á. Löve & D. Löve// Subspecies: Cyanus triumfettii subsp. axillaris (Čelak.) Štěpánek

Determined according to the botanical keys provided by Dostál in Tutin et al. (1976) as Centaurea triumfettii subsp. aligera (Gugler) Dostál, Bot. Jour. Linn. Soc. 71: 208. (1976) (C. variegata var. aligera Gugler) and Gajić in Josifović (1975) as Centaurea triumfettii var. axillaris (Willd.) Hayek (Centaurea axillaris Willd.). The accepted name in the Euro+Med PlantBase (Greuter, 2006+) is Cyanus triumfettii subsp. axillaris (Čelak.) Štěpánek in Slavík & Štěpánková, Květena České Republ. 7: 455. 2004.

Lifeform: Perrenial; hemicryptophyte according to the Raunkiær system.

Distribution: Native from Southeast Europe to Southwest Siberia and Iran according to the distribution information provided by the Plants of the World Online portal (Royal Botanic Gardens, Kew).

Habitat:  Edges and clearings of oak forests.

Flowering period: Summer: JUN.

Cyanus triumfettii subsp. axillaris – habitat. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – habit. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – habit. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – habit. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – habit. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stems with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stem with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stem with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stem with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stem with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper branched part of stem with capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitula. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper part of stem with capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – florets. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – florets. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – florets. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – florets. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – florets. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – involucral bracts. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – involucral bracts. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – involucral bracts. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – peduncle. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – peduncle. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – fruiting capitulum. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – upper cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with lower cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with lower cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – stem with lower cauline leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – winged stem. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – winged stem. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – winged stem. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – winged stem. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – lower part of stem and petiole of lower cauline leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – adaxial side of cauline leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – adaxial side of cauline leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – abaxial side of cauline leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – abaxial side of cauline leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaves. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – basal leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – adaxial side of basal leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – abaxial side of basal leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – adaxial side of basal leaf. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.
Cyanus triumfettii subsp. axillaris – root. Osogovo, between the villages of Pantelej and Rajchani, 21.06.2023. Photo by Trajche Dimitrovski.

References:

Centaurea axillaris Willd. Plants of the World Online portal, Royal Botanic Gardens, Kew. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:189981-1 [Accessed 24.06.2023].

Centaurea L. Dostál, J. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1976). Flora Europaea. Volume 4: Plantaginaceae to Compositae (and Rubiaceae). Cambridge: Cambridge University Press, pp. 254 – 301.

Cyanus triumfettii subsp. axillaris. Greuter, W. (2006+): Compositae (pro parte majore). – In: Greuter, W. & Raab-Straube, E. von (ed.): Compositae. Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet. https://europlusmed.org/cdm_dataportal/taxon/1caf816b-1dbb-4b43-9db4-8f36a8573492 [Accessed 24.06.2023].

Rod Centaurea L. Gajić, M. in Josifović (1975). Flora SR Srbije VII. Srpska Akademija Nauka i Umetnosti, Beograd, pp. 229 – 264.

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