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Ver. 19.2
April 17th, 2024

490.0000 Family Boraginaceae Juss.; Gen. Pl.: 128 (1789), nom. cons., s. str.
118 genera, ca. 2000 spp.
Species listed in database: 2033; hybrids: 45
= Anchusaceae Vest; Anleit. Stud. Bot.: 274, 302 (1818)
= Cerinthaceae Bercht. & J.Presl; Prir. Rostlin: 244 (1820)
= Cynoglossaceae Döll; Rhein. Fl.: 406 (1843)
= Echiaceae Raf.; Fl. Tellur. 2: 61 (1837)
= Onosmaceae Martinov; Tekhno-Bot. Slovar: 437 (1820)
490.3527 Genus Lappula Moench; Methodus: 416 (1794)
Distribution: temp. Eurasia (most), some Mediterranean, few W-North America and disjunct temperate South America
Species listed in database: 82
= Cynoglossospermum Siegesb. ex Kuntze; Revis. Gen. Pl. 3: II. 204 (1898)
= Echinospermum Sw. ex Lehm.; Pl. Asperif. Nucif.: 113 (1818)
= Omphalolappula Brand; Engl. Pflanzenreich, Borag.-Borraginoid.-Cryptanth.135 (1931)
= Sclerocaryopsis Brand; Engl. Pflanzenreich, Borag.-Borraginoid.-Cryptanth. 98 (1931)
= Tianschaniella B.Fedtsch. ex Popov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 337 (1951)
species :
Lappula aktaviensis Popov & Zakirov; Kom., Fl. URSS 19: 712 (1953)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Lappula alaica (Popov) Nabiev; Opred. Rast. Sred. Azii 8: 138 (1986)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Lepechiniella alaica Popov; Kom., Fl. URSS 19: 716 (1953)
species :
Lappula albiflora (Riedl) Khoshsokhan & Kaz.Osaloo; Ann. Bot. Fenn. 55(4-6): 303 (2018)
Distribution: Iran
34 IRN
Synonyms: hidden
= Lepechiniella albiflora Riedl; Rech.f., Fl. Iranica 48, 83 (1967)
species :
Lappula anocarpa C.J.Wang; Bull. Bot. Res., Harbin l(4): 93 (1981)
Distribution: China (C-Gansu, NE-Gansu)
36 CHI
species :
Lappula badachschanica (Popov) Popov; Kom., Fl. URSS 19: 431 (1953)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Lappula semiglabra var. badachschanica Popov
species :
Lappula baitenovii Kudab.; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Kaz. SSR 13: 58 (1983)
Distribution: Kazakhstan
32 KAZ
species :
Lappula balchaschensis Popov ex Golosk.; Fl. Rast. Res. Kaz.: 73 (1975)
Distribution: Siberia (Altai, Tuva); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang); Mongolia
30 SIB 32 KAZ KGZ TZK UZB 36 CHI 37 MON
Synonyms: hidden
= Lappula balchaschensis Popov ex Pavlov; Bot. J., URSS 30: 190 (1945), in clavi, sine descr. lat
species :
Lappula barbata (M.Bieb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. IV(3a): 107 (1897)
Distribution: Hungary; C- & NE-Spain; North Macedonia; Albania; Romania; Bulgaria; Greece; E-European Russia; Ukraine; Crimea; Morocco; Algeria; Turkmenistan; Northern Caucasus; Georgia [Caucasus]; Armenia; ?Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia, SSW-Anatolia, W-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Lebanon (Antilebanon); Jordania (S-Jordania, W-Jordania)
11 HUN 12 SPA 13 ALB BUL GRC NMA ROM 14 KRY RUS UKR 20 ALG MOR 32 TKM 33 ARM GEO NCS 34 IRN JOR LEB TUR
Synonyms: hidden
= Anchusa leucojifolia Lehm.; Pl. Asperif. Nucif. 1: 249 (1818)
= Cynoglossospermum barbatum f. zapateri Pau; Bol. Soc. Esp. Hist. Nat. 1: 154 (1901)
= Echinospermum barbatum (M.Bieb.) Lehm.; Pl. Asperif. Nucif.: 128 (1818)
= Echinospermum barbatum subsp. aragonense Rev. & Freyn; Willk., Suppl. Prodr. Fl. Hispan.: 166 (1893)
= Echinospermum barbatum var. brevistylum Ball; J. Linn. Soc., Bot. 16: 570 (1878)
= Echinospermum barbatum var. cariense (Boiss.) Boiss.; Fl. Orient. 4: 250 (1879)
= Echinospermum myosotiflorum A.DC.; Prodr. [A. DC.] 10: 141 (1846)
= Echinospermum triseriale Bunge; Index Seminum (TU, Dorpatensis) 1840: 8 (1840)
= Echinospermum triseriale Kar. & Kir. ex Ledeb.; Fl. Ross. 3: 162 (1847)
= Echinospermum zapateri Pau ex Magnier; Scrin. fl. select. 13: 72 (1895)
= Lappula barbata subsp. aragonense (E.Rev. & Freyn ex Willk.) Mateo; Cat. Flor. Prov. Teruel: 46 (1990)
= Lappula barbata var. brevistyla (Ball) Murb.; Contr. Fl. Maroc [Murbeck] 2: 22 (1923)
= Lappula echinata var. barbata (M.Bieb.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 214 (1887)
= Lappula saxatilis (Pall.) Kusn.; Fl. Caucasus Crit. 4: II. 183 (1913)
= Lappula zapateri (Pau) O.Bolòs & Vigo; Collect. Bot. (Barcelona) 11: 56 (1979)
= Myosotis barbata M.Bieb.; Fl. Taur.-Caucas. 1: 121 (1808)
= Myosotis saxatilis Pall.; Nova Acta Acad. Sci. Imp. Petrop. Hist. Acad. 1792: 306 (1792), nom. nud.
= Rochelia barbata (M.Bieb.) Roem. & Schult.; Syst. Veg. ed. 15[bis]. 4: 110 (1819)
variant :
var. cariensis (Boiss.) Brand; Pflanzenr. IV. 252: 141 (1931)
Distribution: Tajikistan; Iran; Pakistan (Baluchistan, Quetta, Waziristan, Kurram, Chitral, Astor); NW-India (Himachal Pradesh, Jammu & Kashmir, Punjab, Uttar Pradesh, Uttarakhand)
32 TZK 34 IRN 40 IND PAK
Synonyms: hidden
= Lappula echinata var. cariensis (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 214 (1887)
= Echinospermum cariense Boiss.; Diagn. Ser. I. 4: 40 (1844)
species :
Lappula botschantzevii Ovczinnikova; Phytotaxa 522(1): 48 (2021)
Distribution: Morocco; Algeria; Tunisia
20 ALG MOR TUN
species :
Lappula brachycentra (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang)
32 KAZ KGZ TKM TZK UZB 36 CHI
Synonyms: hidden
= Cynoglossospermum brachycentrum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum brachycentrum Ledeb.; Fl. Altaic. 1: 203 (1829)
= Echinospermum polymorphum var. tuberculatum Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 26: 550 (1910)
= Hackelia brachycentra (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Lappula tuberculata (Lipsky) Zakirov; Trudy Uzbeksk. Gosud. Univ., n.s., 28(15): 18 (1941)
species :
Lappula brachycentroides Popov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 314 (1951)
Distribution: Siberia (Altai); Kazakhstan
30 SIB 32 KAZ
species :
Lappula caespitosa C.J.Wang; Acta Phytotax. Sin. 18: 521 (1980)
Distribution: SW-Tibet
36 TIB
Remarks: probably a synonym of Lappula alatavica
species :
Lappula campanulata (Popov & Zakirov) Sennikov; Checkl. Vasc. Pl. Tian-Shan 312 (2021)
Distribution: Kazakhstan; Uzbekistan; Tajikistan
32 KAZ TZK UZB
Synonyms: hidden
= Rochelia campanulata Popov & Zakirov; Not. Syst. Herb. Inst. Bot. & Zool. Acad. Sci. Uzbekistan. 10: 10 (1948)
species :
Lappula cardiosepala (Bunge) Sennikov; Checkl. Vasc. Pl. Tian-Shan 312 (2021)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Turkey (SE-Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (EC-Iran, NE-Iran: Mts., W-Iran, Iranian Aserbaijan); Afghanistan (Baghlan, Balkh, Bamyan, Faryab, Kabul, Kunar / Nuristan, Paktia / Khost, Samangan, Takhar); NW-Tibet; Pakistan (Chitral, Swat, Gilgit, Baltistan); NW-India (Himachal Pradesh, Jammu & Kashmir)
32 KAZ KGZ TKM TZK UZB 33 ARM 34 AFG IRN IRQ TUR 36 TIB 40 IND PAK
Synonyms: hidden
= Rochelia cardiosepala Bunge; Mém. Sav. Etr. Acad. Petersb. 7: 420 (1851), et Beitr. Fl. Russl.: 244 (1852)
species :
Lappula cenchroides A.Nelson; Bull. Torrey Bot. Club, 243 (1899)
Name: Great Plains stickseed
Distribution: USA (Connecticut, Massachusetts, Montana, North Dakota, Nebraska, South Dakota, Wyoming)
73 USA 74 USA 75 USA
Synonyms: hidden
= Echinospermum cenchroides (A.Nelson) K.Schum.; Just, Jahresb. 27: I. 522 (1901)
= Lappula cenchrusoides A.Nelson; Bull. Torrey Bot. Club 26: 243 (1899), sphalm.
species :
Lappula coronifera Popov; Kom., Fl. URSS 19: 713 (1953)
Distribution: Kazakhstan; Mongolia
32 KAZ 37 MON
species :
Lappula cynoglossoides (Lam.) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
Distribution: Namibia; South Africa (Free State, N-Cape Prov., W-Cape Prov., E-Cape Prov.); Lesotho
27 LES NAM SAF
Synonyms: hidden
= Cynoglossospermum capense (A.DC.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum cynoglossodes (Lam.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossum echinatum Thunb.; Prodr. Pl. Cap. 84 (1794)
= Echinospermum capense A.DC.; Prodr. [A. DC.] 10: 138 (1846)
= Echinospermum cynoglossoides (Lam.) Lehm.; Asperif. 131 (1818)
= Echinospermum ecklonianum Lehm. ex Brand; Repert. Spec. Nov. Regni Veg. 18: 312 (1922)
= Lappula capensis (A.DC.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1894)
= Lappula eckloniana Brand; Repert. Spec. Nov. Regni Veg. 18: 312 (1922)
= Myosotis cynoglossoides Lam.; Tabl. Encyc. 1: 396 (1791)
= Myosotis echinata (Thunb.) Thunb.; Fl. Cap. 2: 352 (1818)
= Rochelia cynoglossoides (Lam.) Roem. & Schult.; Syst. 4: 111, 783 (1819)
species :
Lappula deserticola C.J.Wang; Bull. Bot. Res., Harbin l(4): 81 (1981)
Distribution: China (Gansu, Nei Mongol)
36 CHI
variant :
var. longicalyx X.M.Pi & Z.J.Wang; J. Xinjiang Univ. (Nat. Sci. Ed.) 14(2): 67 (1997)
Distribution: China (Xinjiang)
36 CHI
species :
Lappula dichotoma Charit.; Dopoln. Fl. i Rastitel´n. Yuga Zapadnoi Sibiri 170 (-171) (2020)
Distribution: Siberia (W-Siberia)
30 SIB
Remarks: provisionally accepted name, will highly likely turn out to be a synonym of a more widespread species
species :
Lappula diploloma (Fisch. & C.A.Mey.) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
Distribution: Kazakhstan; Mongolia
32 KAZ 37 MON
Synonyms: hidden
= Cynoglossospermum diploloma (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum diploloma Fisch. & C.A.Mey.; Enum. Pl. Nov. 1: 36 (1841)
= Echinospermum montanum (Greene) K.Schum.; Just´s Bot. Jahresber. 27(1): 522 (1901)
= Lappula montana Greene; Pittonia 4: 96 (1899)
species :
Lappula duplicicarpa Pavlov; Vestn. Akad. Nauk SSSR 5: 90 (1952)
Distribution: Kazakhstan; Uzbekistan; Tajikistan; China (Qinghai, N-Xinjiang)
32 KAZ TZK UZB 36 CHI
variant :
var. brevispinula C.J.Wang; Bull. Bot. Res., Harbin l(4): 84 (1981)
Distribution: China (N-Xinjiang)
36 CHI
variant :
var. densihispida C.J.Wang; Bull. Bot. Res., Harbin l(4): 83 (1981)
Distribution: China (W-Gansu, N-Xinjiang)
36 CHI
species :
Lappula dzharkentica Popov ex Golosk., Bot.Mater Gerb.Inst.Bot.Akad.Nauk Kaz.SSR 7: 23 (1971)
Distribution: Siberia (Altai, Tuva); Kazakhstan
30 SIB 32 KAZ
Synonyms: hidden
= Lappula semiglabra var. dzharkentica Popov
= Lappula dzharkentica (Popov) Popov; Kom., Fl. URSS 19: 431 (1953)
species :
Lappula fetisowii (Regel) Sennikov; Checkl. Vasc. Pl. Tian-Shan 312 (2021)
Distribution: Kazakhstan; Kyrgyzstan; Tajikistan; China (W-Xinjiang)
32 KAZ KGZ TZK 36 CHI
Synonyms: hidden
= Eritrichium fetisowii var. minor Popov ex Ovczinnikova; Rast. Mir Aziatsk. Rossii 7: 46 (2011)
= Eritrichium fetisowii Regel; Trudy Imp. S.-Peterburgsk., Bot. Sada 6: 340 (1879)
species :
Lappula fruticulosa Ovczinnikova; Bot. Zhurn. (Moscow & Leningrad) 86(5): 133 (2001)
Distribution: Siberia (Altai)
30 SIB
species :
Lappula glabrata Popov ex Pavlov; Bot. J., URSS 30: 191 (1945), in clavi, sine descr. lat., Popov in Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 322 (1951)
Distribution: Kazakhstan
32 KAZ
Conservation Status: IUCN R
species :
Lappula granulata (Krylov) Popov; Kom., Fl. URSS-19: 426 (1953)
Distribution: China (Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Ningxia, Shaanxi, Shandong, Shanxi); Mongolia
36 CHI 37 MON
Synonyms: hidden
= Lappula gansuensis X.D.Wang & C.J.Wang; Acta Phytotax. Sin. 30(4): 369 (1992)
= Lappula marginata var. granulata Krylov; Fl. Zap./West Sib. 9: 2248 (1937)
= Lappula platyacantha W.T.Wang ex C.J.Wang; Bull. Bot. Res. North-East. Forest. Inst. 1(4): 89 (1981)
species :
Lappula heteracantha (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. IV(3a): 107 (1897)
Distribution: Czech Republic [I]; Slovakia [I]; Hungary [I]; France [I]; Italy [I]; Croatia [I]; European Russia; Crimea; South Africa [I] (widespread); Lesotho [I]; Siberia (Altai, Tuva, W-Siberia); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Iran (NE-Iran: Mts., N-Iran, W-Iran); Afghanistan (Bamyan, Orozgan / Daykundi); Mongolia; North Korea; Pakistan (Gilgit, Astor); NW-India (Dras, Jammu & Kashmir)
11 cze hun slo 12 fra 13 cro ita 14 KRY RUS 27 les saf 30 SIB 32 KAZ 33 GEO NCS 34 AFG IRN 37 MON 38 NKO 40 IND PAK
Synonyms: hidden
= Cynoglossospermum cymosum (Stschegl.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum semicinctum (Steven) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum bornmuelleri Hausskn. ex Bornm.; Beih. Bot. Centralbl. 20(2): 190 (1906)
= Echinospermum cymosum Stschegl.; Bull. Soc. Imp. Naturalistes Moscou 27: I. 184 (1854)
= Echinospermum heteracanthum Ledeb.; Suppl. Ind. Sem. Hort. Dorpat. 1823: 3 (1823)
= Echinospermum semicinctum Steven; Bull. Soc. Imp. Naturalistes Moscou 24(I): 605 (1851)
= Lappula echinata var. heteracantha (Ledeb.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 214 (1887)
= Lappula heteracantha subsp. heterocarpa (Klokov & Artemczuk) Holub; Fl. Slovenska 5(1): 153 (1993)
= Lappula heterocarpa Klokov & Artemczuk; Ukrayin. Bot. Zhurn. 9: No. 3, 83 (1952)
= Lappula myosotis subsp. heteracantha (Ledeb.) Á.Löve & D.Löve
= Lappula semicincta (Steven) Dobrocz.; Bordzil. et al., Fl. URSR 8: 485 (1958), isonym
= Lappula semicincta (Steven) Popov; Kom., Fl. URSS 19: 428 (1953)
= Lappula squarrosa subsp. heteracantha (Ledeb.) Chater; Bot. J. Linn. Soc. 64: 380 (1971)
species :
Lappula heteromorpha C.J.Wang; Bull. Bot. Res., Harbin l(4): 95 (1981)
Distribution: China (Nei Mongol, Shanxi)
36 CHI
species :
Lappula himalayensis C.J.Wang; Acta. Phytotax. Sin. 18: 521 (1980)
Distribution: S-Tibet; India (Jammu & Kashmir)
36 TIB 40 IND
species :
Lappula iljinii Sennikov; Checkl. Vasc. Pl. Tian-Shan 313 (2021), nom. nov.
Distribution: Kyrgyzstan
32 KGZ
Synonyms: hidden
= Eritrichium tianschanicum Iljin ex Ovczinnikova; Bot. Zhurn. (Moscow & Leningrad) 88(7): 79 (2003)
species :
Lappula intermedia (Ledeb.) Popov; Kom., Fl. URSS-19:440 (1953)
Distribution: Siberia (Altai, W-Siberia, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva); Russian Far East (Khabarovsk, Primorye); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, NW-Sichuan, Xinjiang); Tibet; Mongolia; NW-India (Jammu & Kashmir)
30 SIB 31 RFE 32 KAZ KGZ TKM TZK UZB 36 CHI TIB 37 MON 40 IND
Synonyms: hidden
= Cynoglossospermum anisacanthum (Turcz. ex Bunge) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum anisacanthum Turcz. ex Bunge; Index Seminum (TU, Dorpatensis) 1840: 8 (1840)
= Echinospermum anisacanthum Turcz. ex Ledeb.; Fl. Ross. (Ledeb.) 3(1,8): 156 (1847)
= Echinospermum intermedium Ledeb.; Fl. Altaic. 1: 199 (1829)
= Hackelia intermedia (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Lappula anisacantha (Turcz. ex Bunge) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
= Lappula anisacantha var. sibirica Popov ex Ovchinnikova; Rast. Mir Aziatsk. Rossii 16(2): 136 (2023)
= Lappula anisacantha var. uniserialis Popov ex Ovchinnikova; Rast. Mir Aziatsk. Rossii 16(2): 137 (2023)
species :
Lappula karelinii (Fisch. & C.A.Mey.) Kamelin; Novon 3: 263 (1993)
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
Synonyms: hidden
= Echinospermum karelinii Fisch. & C.A.Mey.; Ind. Sem. Hort. Petrop. 11: 67 (1845)
= Echinospermum redowskii var. karelinii (Fisch. & C.A.Mey.) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 84 (1868)
= Lappula xinjiangensis C.Y.Yang ex C.J.Wang; Bull. Bot. Res. North-East. Forest. Inst. 1(4): 84 (1981)
species :
Lappula ketmenica Kudab.; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Kaz. SSR 11: 60 (1979)
Distribution: Kazakhstan
32 KAZ
species :
Lappula korshinskyi Popov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 323 (1951)
Distribution: Kyrgyzstan; Tajikistan; China (SW-Xinjiang)
32 KGZ TZK 36 CHI
Synonyms: hidden
= Lappula ferganensis (Popov) Kamelin & G.L.Chu; G. L. Chu, Kamelin, R. R. Mill & M. G. Gilbert, Novon 5: 17 (1995)
= Lappula platyptera C.J.Wang; Bull. Bot. Res. North-East. Forest. Inst. 1(4): 91 (1981)
= Lepechiniella ferganensis Popov; Kom., Fl. URSS 19: 715 (1953)
species :
Lappula krylovii Ovczinnikova, Pjak & A.L.Ebel; Turczaninowia 7(2): 5 (2004)
Distribution: Siberia (Altai)
30 SIB
species :
Lappula kulikalonica Zakirov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 13: 43 (1950)
Distribution: Siberia (Altai); Uzbekistan; Tajikistan
30 SIB 32 TZK UZB
species :
Lappula kuprijanovii Ovchinnikova; Rast. Mir Aziatsk. Rossii 14(1): 10 (2021)
Distribution: Kazakhstan
32 KAZ
species :
Lappula laxa X.M.Pi & Z.J.Wang; J. Xinjiang Univ. (Nat. Sci. Ed.) 14(2): 66 (1997), nom. illeg., non (G. Don) J. F. Macbr.
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Lappula lenensis Popov ex Ovczinnikova; Fl. Sibir. (Pyrolac.-Lamiac.) 11: 137 (1997)
Distribution: Siberia (Irkutsk, Krasnoyarsk, Yakutia)
30 SIB
species :
Lappula leonardii Riedl; Bull. Jard. Bot. Nation. Belg. 45: 128 (1975)
Distribution: Iran (EC-Iran, S-Iran, SW-Iran)
34 IRN
species :
Lappula lipschitzii Popov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 330 (1951)
Distribution: Kazakhstan
32 KAZ
species :
Lappula lipskyi Popov; Bot. Mater. Gerb. Bot. Inst. Acad. Nauk SSSR 14: 311 (1951)
Distribution: Kazakhstan; China (NW-Xinjiang); W-Mongolia
32 KAZ 36 CHI 37 MON
species :
Lappula macra Popov ex Pavlov; Bot. J., URSS 30: 190 (1945), in clavi, sine descr. lat., Popov in Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 323 (1951), in obs., et Bot. Zhurn. SSSR 30(4): 190 (1954)
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
species :
Lappula macrantha (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
Distribution: Kazakhstan; China (NW-Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
Synonyms: hidden
= Cynoglossospermum macranthum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum macranthum Ledeb.; Fl. Altaic. 1: 205 (1829)
= Hackelia macrantha (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Oreogenia ferghanica Brand; Repert. Spec. Nov. Regni Veg. 22: 103 (1925)
species :
Lappula marginata (M.Bieb.) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
Name: Margined stickseed
Distribution: Hungary [I]; North Macedonia [I]; S-European Russia; E-European Russia; Siberia (Altai, Krasnoyarsk, Tuva); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Armenia; Iran (NE-Iran: Mts., N-Iran); Afghanistan (Helmand, Herat, Kandahar); China (Gansu, Qinghai, Xinjiang); Mongolia; Pakistan (Makran, Baltistan); NW-India (Jammu & Kashmir); USA [I] (Colorado [I], Oregon [I], Pennsylvania [I])
11 hun 13 nma 14 RUS 30 SIB 32 KAZ KGZ TKM TZK UZB 33 ARM NCS 34 AFG IRN 36 CHI 37 MON 40 IND PAK 73 usa 75 usa
Remarks: often treated under the name L. semiglabra which belongs to L. patula
Synonyms: hidden
= Cynoglossospermum marginatum (M.Bieb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum marginatum (M.Bieb.) Lehm.; Asperif. 130 (1818)
= Echinospermum patulum var. pterocarpum Hochr.; Annuaire Conserv. Jard. Bot. Genève 7-8: 171 (1904)
= Echinospermum semiglabrum var. bungei Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 87 (1868)
= Echinospermum semiglabrum var. karelinii Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 87 (1868)
= Lappula cymosa B.Fedtsch.; Rastit. Turkest. 663 (1915)
= Lappula semiglabra var. heterocaryoides Popov ex C.J.Wang; Bull. Bot. Res., Harbin 1(4): 83 (1981)
= Myosotis marginata M.Bieb.; Fl. Taur. Caucasus 1: 120 (1808)
= Rochelia marginata (M.Bieb.) Roem. & Schult.; Syst. 4: 110 (1819)
species :
Lappula microcarpa (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfarn. 4(3a): 107 (1897)
Distribution: Siberia (Altai, Buryatia [I], W-Siberia); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Turkey (E-Anatolia, N-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Badakshan, Wakhan, Badghys, Baghlan, Balkh, Bamyan, Faryab, Ghazni, Herat, Kabul, Kandahar, Kunar / Nuristan, Logar, Wardak, Nangarhar, Orozgan / Daykundi, Paktia / Khost, Parwan, Samangan, Takhar, Zabul); China (N-Xinjiang); SE-Tibet; S-Tibet; Mongolia; Pakistan (Kurram, Baluchistan, Quetta, Chitral, Baltistan); Nepal; NW-India (Himachal Pradesh, Jammu & Kashmir)
30 SIB 32 KAZ KGZ TKM TZK UZB 34 AFG IRN IRQ TUR 36 CHI TIB 37 MON 40 IND NEP PAK
Synonyms: hidden
= Cynoglossospermum microcarpum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum oligacanthum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum filiforme Godet ex A.DC.; Prodr. [A. DC.] 10: 140 (1846)
= Echinospermum microcarpum Ledeb.; Fl. Alt. 1: 202 (1829)
= Echinospermum oligacanthum Ledeb.; Fl. Ross. 3: 161 (1846-51)
= Echinospermum polymorphum Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 26: 541 (1910)
= Echinospermum polymorphum var. aculeatum Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 26: 547 (1910)
= Echinospermum polymorphum var. minimum Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 26: 551 (1910)
= Echinospermum redowskii var. oligacanthum (Ledeb.) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 84 (1868)
= Echinospermum rigidum A.DC.; Prodr. [A. DC.] 10: 141 (1846)
= Echinospermum stylosum Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 715 (1841)
= Hackelia microcarpa (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Lappula filiformis Kusn.; Fl. Caucasus Crit. 4: II. 197 (1913)
= Lappula microcarpa var. heterogenea X.D.Wang & C.J.Wang; Acta Phytotax. Sin. 30(4): 369 (1992)
= Lappula microcarpa var. micrantha Bornm.; Bot. Jahrb. Syst. 66(2): 235 (1934)
= Lappula polymorpha (Lipsky) B.Fedtsch.; Bot. Explor. Siberia & Turkestan 1911: 88 (1912)
= Lappula rigida (A.DC.) Popov; Kom., Fl. URSS 19: 464 (1953)
species :
Lappula mogoltavica Popov ex Czukav.; Izv. Akad. Nauk Tadzh. SSR, Biol. Nauk, 3(92): 92 (1983)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
species :
Lappula monocarpa C.J.Wang; Bull. Bot. Res., Harbin l(4):98 (1981)
Distribution: China (NW-Xinjiang)
36 CHI
species :
Lappula nevskii Raenko; Bot. Zhurn. 72(1): 99 (1987)
Distribution: Turkmenistan
32 TKM
species :
Lappula nuratavica Nabiev & Zakirov; Fl. Uzbekist. 5: 218, 631 (1961)
Distribution: Uzbekistan
32 UZB
species :
Lappula occidentalis (S.Watson) Greene; Pittonia 4: 97 (1901)
Name: Flatspine stickseed
Distribution: Alaska; Canada (Alberta, British Columbia, Manitoba, Northern Territories, Saskatchewan, Yukon); USA (Arkansas, Arizona, California, Colorado, Connecticut, Iowa, Idaho, Illinois, Kansas, Massachusetts, Maine, Minnesota, Missouri, Montana, North Dakota, Nebraska, New Mexico, Nevada, New York, Oklahoma, Oregon, Pennsylvania, South Carolina, South Dakota, Texas, Utah, Washington State, Wisconsin, Wyoming); Mexico (Sonora, Chihuahua, Coahuila); Argentina (Chubut, La Pampa, La Rioja, Mendoza, Neuquen, Rio Negro, Santa Cruz, San Juan, San Luis); Chile (Magallanes)
70 ASK CDN 71 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA 79 MEX 85 ARG CLE
Synonyms: hidden
= Echinospermum brachycentrum var. brachystylum A.Gray; Proc. Amer. Acad. Arts 21: 413 (1886)
= Echinospermum desertorum (Greene) K.Schum.; Just´s Bot. Jahresber. 27(1): 522 (1901)
= Echinospermum occidentale (S.Watson) K.Schum.; Just´s Bot. Jahresber. 27, Abt. 1: 522 (1901)
= Echinospermum patagonicum Speg.; Anal. Soc. Ci. Argentina. 53: 79 (1902)
= Echinospermum redowskii var. occidentale S.Watson; Botany [Fortieth Parallel] 246 (-247
= Lappula brachystyla (A.Gray) J.F.Macbr.; Contr. Gray Herb. 48: 40 (1916)
= Lappula desertorum Greene; Pittonia 4: 95 (1899)
= Lappula echinata f. brachystyla (A.Gray) B.Boivin; Naturaliste Canad. 93: 1060 (1967)
= Lappula echinata var. occidentalis (S.Watson) B.Boivin; Naturaliste Canad. 93: 1060 (1967)
= Lappula leucotricha Rydb.; Bull. Torrey Bot. Club 36: 676 (1909)
= Lappula patagonica (Speg.) Macloskie; Rep. Princeton Univ. Exped. Patagonia, Botany 8(Suppl.): 214 (1915)
= Lappula redowskii f. brachystyla (A.Gray) Scoggan; Fl. Canada 1: 52 (1978)
= Lappula redowskii subsp. occidentalis (S.Watson) Á.Löve & D.Löve; Taxon 31(2): 353 (1982)
= Lappula redowskii var. desertorum (Greene) I.M.Johnst.; Contr. Arnold Arbor. 3: 93 (1932)
= Lappula redowskii var. occidentalis (S.Watson) Rydb.; Contr. U. S. Natl. Herb. 3(3): 170 (1895)
variant :
var. cupulata (A.Gray) L.C.Higgins; Brigham Young Univ. Sci. Bull., Biol. 16(3): 62 (1972)
Name: Flatspine stickseed
Distribution: Canada (Alberta, British Columbia, Saskatchewan); USA (Arizona, California, Colorado, Iowa, Idaho, Kansas, Missouri, Montana, North Dakota, Nebraska, New Mexico, Nevada, Oklahoma, Oregon, South Dakota, Texas, Utah, Washington State, Wyoming); Mexico (Coahuila, Sonora)
71 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Echinospermum collinum (Greene) K.Schum.; Just´s Bot. Jahresber. 27(1): 522 (1901)
= Echinospermum coronatum (Greene) K.Schum.; Just´s Bot. Jahresber. 27(1): 522 (1901)
= Echinospermum heterospermum (Greene) K.Schum.; Just´s Bot. Jahresber. 27(1): 522 (1901)
= Echinospermum redowskii var. cupulatum A.Gray; Bot. California [W. H. Brewer] 1: 530 (1876), nom. illeg. superfl.
= Echinospermum scabrosum Buckley; Proc. Acad. Nat. Sci. Philadelphia 13: 462 (1862)
= Echinospermum texanum Scheele; Linnaea 25: 260 (1852)
= Lappula collina Greene; Pittonia 4: 96 (1899)
= Lappula columbiana A.Nelson; Bot. Gaz. 34: 28 (1903)
= Lappula coronata Greene; Pittonia 4: 94 (1899)
= Lappula cupulata (A.Gray) Rydb.; Bull. Torrey Bot. Club, 31 (1901)
= Lappula cupulata var. foliosa (A.Nelson) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 40 (1916)
= Lappula desertorum var. foliosa A.Nelson; Bull. Torrey Bot. Club 27: 267 (1900)
= Lappula echinata f. cupulata (A.Gray) B.Boivin; Naturaliste Canad. 93: 1060 (1967)
= Lappula foliosa (A.Nelson) A.Nelson; Coult. & Nels., Man. Bot. Rocky Mt., ed. 2: 413 (1909)
= Lappula heterosperma Greene; Pittonia 4: 94 (1901)
= Lappula heterosperma var. homosperma A.Nelson; Bot. Gaz. 34: 29 (1902)
= Lappula redowskii f. cupulata (A.Gray) Scoggan; Fl. Canada 1: 52 (1978)
= Lappula redowskii var. cupulata (A.Gray) M.E.Jones; Bull. Montana Univ., Biol. Ser. ser. 15: 44 (1910)
= Lappula redowskii var. texana (Scheele) Brand; Pflanzenr. (Engl.) [Heft 97] 4, Fam. 252: 150 (1931)
= Lappula texana (Scheele) Britton; Mem. Torrey Bot. Club 5: 273 (1894)
= Lappula texana var. columbiana (A.Nelson) I.M.Johnst.; Contr. Gray Herb. 70: 50 (1924)
= Lappula texana var. coronata (Greene) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 41 (1916)
= Lappula texana var. foliosa (A.Nelson) I.M.Johnst.; Contr. Gray Herb. 70: 51 (1924)
= Lappula texana var. heterosperma (Greene) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 41 (1916)
= Lappula texana var. homosperma (A.Nelson) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 41 (1916)
species :
Lappula occultata Popov; Bot Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 14: 331 (1951)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Afghanistan (Bamyan); China (Xinjiang); Mongolia
32 KAZ KGZ TKM TZK UZB 34 AFG 36 CHI 37 MON
Synonyms: hidden
= Lappula laevimarginata Riedl; Rech.f., Fl. Iranica 48, 78 (1967)
= Lappula occultata var. heterocaryoides X.M.Pi & Z.J.Wang; J. Xinjiang Univ. (Nat. Sci. Ed.) 14: 67 (1997)
= Lappula sinaica var. occultata (Popov) Pavlov; Bot. Zhurn. S.S.S.R. 30(4): 191 (1945)
species :
Lappula parvula Nabiev & Zakirov; Fl. Uzbekist. 5: 222, 632 (1961)
Distribution: Uzbekistan
32 UZB
species :
Lappula patula (Lehm.) Menyh.; Kal. Vid. Növén. 120 (1877)
Name: Abstehender Igelsame (DE)
Distribution: Germany [I]; Switzerland [I]; Austria [I]; Slovakia; C- & NE-Spain; France; Italy; Sicily; Croatia; Romania; Bulgaria; Greece; Belarus [I]; Crimea; Morocco; Algeria; Tunisia; Siberia (Altai); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; ?Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Baghlan, Bamyan); China (N-Xinjiang); Mongolia; NW-India (Ladakh, Dras, Jammu & Kashmir)
11 aut ger SLO swi 12 FRA SPA 13 BUL CRO GRC ITA ROM SIC 14 blr KRY 20 ALG MOR TUN 30 SIB 32 KAZ KGZ TKM TZK UZB 33 ARM GEO NCS 34 AFG IRN TUR 36 CHI 37 MON 40 IND
Synonyms: hidden
= Cynoglossospermum caspium (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 2: 437 (1891), as ´caspicum´
= Cynoglossospermum patulum (Lehm.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum semiglabrum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossum patulum (Lehm.) E.H.L.Krause; Sturn. Fl Deutschland, ed. 2, 11: 33 (1903)
= Echinospermum brachysepalum Claus; Beitr. Pflanzenk. Russ. Reiches 8: 240 (1851)
= Echinospermum caspium Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 5: 34 (1839)
= Echinospermum patulum Lehm.; Pl. Asperif. Nucif.: 124 (1818)
= Echinospermum semiglabrum Ledeb.; Fl. Altaic. 204 (1829)
= Echinospermum skorpili Velen.; Fl. Bulg., Suppl. 1: 206 (1898)
= Echinospermum squarrosum (M.Bieb.) Link; Enum. Hort. Berol. 50: 165 (1821)
= Hackelia patula (Lehm.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Hackelia semiglabra (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Lappula caspia (Fisch. & C.A.Mey.) Czerep.; Sosud. Rast. SSSR: 114 (1981), isonym
= Lappula caspia (Fisch. & C.A.Mey.) Popov; Kom., Fl. URSS 19: 431 (1953), ex Dobrocz.
= Lappula marginata subsp. patula (Lehm.) Soó; Acta Bot. Acad. Sci. Hung., 23(3-4): 382 (1977 publ. 1978)
= Lappula patula (Lehm.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. IV(3a): 107 (1897)
= Lappula redowskii subvar. semiglabra (Ledeb.) Brand; Engl., Pflanzenr., IV, 252(97): 149 (1931)
= Lappula redowskii subsp. patula (Lehm.) Maire; Jahand. & Maire, Cat. Pl. Maroc: 595 (1934)
= Lappula redowskii var. marginata Maire; Bull. Soc. Hist. Nat. Afrique N. 32: 214 (1941)
= Lappula redowskii var. patula (Lehm.) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 39 (1916)
= Lappula redowskyi subsp. patula (Lehm.) Maire; Jahand. et al., Cat. Pl. Maroc 3: ? (1934)
= Lappula semiglabra (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
= Lappula semiglabra var. subsemiglabra Popov
= Lappula skorpili (Velen.) Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 1916: 630 (1917)
= Lappula subsemiglabra (Popov) Popov; Kom., Fl. URSS 19: 432 (1953)
= Lithospermum patulum (Lehm.) Gand.; Contrib. Fl. Terr. Slav. Merid. 1: 22 (1883)
= Myosotis squarrosa M.Bieb.; Fl. Taur.-Caucasus 1: 120 (1808) (non Retz. (1781))
= Rochelia patula (Lehm.) Roem. & Schult.; Syst. 4: 781 (1819)
= Rochelia squarrosa (M.Bieb.) Roem. & Schult.; Syst. 4: 111, 109 (1819)
species :
Lappula paulsenii (Brand) Popov; Kom., Fl. URSS 19: 464 (1953)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Oreogenia paulsenii Brand; Repert. Spec. Nov. Regni Veg. 22: 103 (1925)
species :
Lappula pavlovii Golosk.; Fl. Rast. Res. Kaz.: 71 (1975)
Distribution: Kazakhstan
32 KAZ
species :
Lappula physacantha Golosk.; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk. Kazahsk. SSR 10: 34 (1977)
Distribution: Siberia (Altai); Kazakhstan; Kyrgyzstan; China (Xinjiang)
30 SIB 32 KAZ KGZ 36 CHI
species :
Lappula popovii Zak.; Not. Syst. Herb. Inst. Bot. & Zool. Acad. Sci. Uzbekistan. 10: 7 (1948)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Lappula pratensis C.J.Wang; Bull. Bot. Res., Harbin l(4): 87 (1981)
Distribution: China (Xinjiang)
36 CHI
species :
Lappula ramulosa C.J.Wang & X.D.Wang; Acta Phytotax. Sin. 30: 367 (1992)
Distribution: China (Xinjiang)
36 CHI
species :
Lappula rechingeri Riedl; Ann. Naturhist. Mus. Wien, 98B(Suppl.): 80 (1996)
Distribution: Pakistan
40 PAK
species :
Lappula redowskii (Hornem.) Greene; Pittonia 2: 182 (1891)
Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Yakutia); Russian Far East (Magadan); Mongolia
30 SIB 31 RFE 37 MON
Synonyms: hidden
= Cryptantha lappula Brand; Repert. Spec. Nov. Regni Veg. 24: 56 (1927)
= Cynoglossospermum affine (Kar. & Kir.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum redowskii (Hornem.) Kuntze; Revis. Gen. Pl. 2: 436 (1891)
= Cynoglossum pilosum Nutt.; Gen. N. Amer. Pl. [Nutt.]. 1: 114 (1818)
= Echinospermum affine Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 14: 714 (1841)
= Echinospermum pilosum Buckley; Proc. Acad. Nat. Sci. Philadelphia 13: 462 (1862)
= Echinospermum redowskii (Hornem.) Lehm.; Asperif. 127 (1818)
= Echinospermum redowskii var. affine (Kar. & Kir.) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 84 (1868)
= Echinospermum redowskii var. strictum (Nees) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 84 (1868)
= Echinospermum strictum Nees; Wied, Reise Nord-America 2: 443 (1841)
= Lappula inconspicua Brand; Repert. Spec. Nov. Regni Veg. 26: 171 (1929)
= Lappula pilosa (Buckley) Hitchc.; Spring Fl. Manhattan 30 (1894)
= Lappula redowskii (Hornem.) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
= Lappula redowskii var. pilosum MacMill.; Metasp. Minnesota Valley: 441 (1892)
= Myosotis redowskii Hornem.; Hort. Hafn. 1: 174 (1813)
= Rochelia redowskii (Hornem.) Roem. & Schult.; Syst. 4: 110. 782 (1819)
species :
Lappula rupestris (Schrenk) Gürke; Engl. & Prantl, Natürl. Pflanzenfam. 4: 3a 107 (1893)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Echinospermum rupestre Schrenk; Bull. PhysMath. Acad. Petersb. 2: 194 (1842)
= Echinospermum microcarpum var. rupestre (Schrenk) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 85 (1868)
species :
Lappula rupicola Zakirov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 13: 41 (1950)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Lappula saissanica Aralbaev; Izv. Akad. Nauk Resp. Kazakhstan, Ser. Biol. Med., 1997(2): 58 (1997)
Distribution: Kazakhstan
32 KAZ
species :
Lappula saphronovae Kamelin; Turczaninowia 2(3): 5 (1999)
Distribution: Kazakhstan
32 KAZ
species :
Lappula semialata Popov; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 327 (1951)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Lappula semnanensis Riedl & Iranshahr; Linz. Biol. Beitr. 17(2): 327 (1985)
Distribution: Iran
34 IRN
species :
Lappula sericata Popov; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 14: 320 (1951)
Distribution: Kazakhstan; Uzbekistan; Tajikistan; China (Xinjiang)
32 KAZ TZK UZB 36 CHI
species :
Lappula shanhsiensis Kitag.; Acta Phytotax. Geobot. 20: 48 (1962)
Distribution: China (Gansu, Hebei, Nei Mongol, Shanxi); Tibet
36 CHI TIB
species :
Lappula spinocarpos (Forssk.) Asch.; Verh. Bot. Vereins Prov. Brandenburg 16, Sitzungsber.: 88 (1874)
subspecies :
subsp. spinocarpos
Distribution: S-European Russia; Morocco; Algeria; Tunisia; Libya; Egypt (Great Southwestern Desert, NE-Egypt, NW-coastal Egypt, SE-Egypt); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia); Iraq (NW-Iraq, SE-Iraq: Mesopotamia, S-Iraq, W-Iraq: Desert); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Bamyan, Farah, Ghazni, Helmand, Herat, Kabul, Kandahar, Logar, Wardak, Nangarhar, Paktia / Khost); Lebanon (Antilebanon); Syria (W-Syrian Mountains); Israel (Rift Valley, N-Negev Desert, Judean Desert, S-Negev Desert, Negev Desert); Sinai peninsula (N-Sinai, N-Sinai, S-Sinai); Jordania (S-Jordania, W-Jordania); Saudi Arabia (C-Saudi Arabia, Dahana, Asir); Kuwait; United Arab Emirates; Oman (Mascat & Oman); China (Xinjiang: Zhungeer basin); Pakistan (Quetta)
14 RUS 20 ALG EGY LBY MOR TUN 32 KAZ KGZ TKM TZK UZB 33 ARM GEO 34 AFG IRN IRQ ISR JOR LEB SIN SYR TUR 35 KUW OMA SAU UAE 36 CHI 40 PAK
Synonyms: hidden
= Anchusa spinocarpos Forssk.; Fl. Aegypt-Arab.: 41 (1775)
= Cynoglossospermum spinocarpum (Forssk.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum spinocarpos (Forssk.) Boiss.; Fl. Orient. 4: (1875)
= Echinospermum tuberculosum Ledeb.; Eichwald, Pl. Nov. 2(1): 6 (1835)
= Echinospermum vahlianum Lehm.; Asperif. 132 (1818)
= Lappula spinocarpa (Forssk.) Asch. ex Kuntze; Act. Hort. Petrop. 10: 215 (1887)
= Lappula spinocarpos var. inermis Botsch.; Bull. Acad. Sci. SSSR, Bot. 1: 364 (1964)
= Lappula tuberculosa Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1894)
= Lappula vahliana (Lehm.) Nábelek; Publ. Fac. Sci. Univ. Masaryk, Brno, No. 70, 23 (1926)
= Myosotis spinocarpos (Forssk.) Vahl; Symb. Bot. 2: 32 (1791)
= Rochelia spinocarpos (Forssk.) Roem. & Schult.; Syst. 4: 111, 783 (1819)
= Sclerocaryopsis spinocarpos (Forssk.) Brand; Engl., Pflanzenr., IV, 252(79): 98 (1931)
subspecies :
subsp. ceratophora (Popov) Y.J.Nasir; Fl. Pakistan, 191: 139 (1989)
Distribution: Turkmenistan; Uzbekistan; Tajikistan; Iraq (SE-Iraq: Mesopotamia); Iran (EC-Iran, NE-Iran: Mts., S-Iran, W-Iran); Afghanistan (Farah, Helmand, Kandahar, Samangan); Pakistan (Baluchistan, Quetta)
32 TKM TZK UZB 34 AFG IRN IRQ 40 PAK
Synonyms: hidden
= Echinospermum ceratophorum Popov; Korovin, Kult. & Popov, Descr. Pl. Nov. Turk. 67, t. 17 (1916)
= Lappula ceratophora (Popov) Popov; Kom., Fl. URSS 19: 417 (1953)
= Sclerocaryopsis ceratophora (Popov) Sadat; Mitt. Bot. Staatssamml. München 28: 22 (1989)
species :
Lappula squarrosa (Retz.) Dumort.; Fl. Belg.: 40 (1827)
Name: Kletten-Igelsame (DE), European stickseed (EN)
Distribution: Denmark [I]; Norway [I]; Sweden; Finland [I]; Netherlands [I]; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, +Hessen, Hessen [I], Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen [I], Rheinland-Pfalz, Schleswig-Holstein, Saarland [I], Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; ?Liechtenstein; Austria; Poland [I]; Czech Republic; Slovakia; Hungary; C- & NE-Spain; Andorra; Baleares; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (NW- & N-Greece, rare N-Peloponnisos, Athos); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; Ukraine; Crimea; ?Algeria; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Sakhalin); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, W-Iran); Afghanistan (Wakhan); China (Gansu, Hebei, W-Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Xinjiang); Mongolia; North Korea; Pakistan (Gilgit, Kurram, Baluchistan, Quetta); NW-India (Jammu & Kashmir); Alaska [I]; Canada [I] (Alberta [I], British Columbia [I], Labrador [I], Manitoba [I], New Brunswick [I], Newfoundland [I], Nova Scotia [I], Northern Territories [I], Ontario [I], Prince Edward Isl. [I], Québec [I], Saskatchewan [I], Yukon [I]); USA [I] (Arkansas [I], Arizona [I], California [I], Colorado [I], Connecticut [I], District of Columbia [I], Iowa [I], Idaho [I], Illinois [I], Indiana [I], Kansas [I], Kentucky [I], Massachusetts [I], Maryland [I], Maine [I], Michigan [I], Minnesota [I], Missouri [I], Montana [I], North Dakota [I], Nebraska [I], New Hampshire [I], New Jersey [I], New Mexico [I], Nevada [I], New York [I], Ohio [I], Oregon [I], Pennsylvania [I], Rhode Island [I], South Dakota [I], Tennessee [I], Texas [I], Utah [I], Virginia [I], Vermont [I], Washington State [I], Wisconsin [I], West Virginia [I], Wyoming [I]); Venezuela [I]
10 den fin nor SWE 11 AUT CZE GER HUN net pol SLO SWI 12 ADO BAL FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 31 RFE 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO NCS 34 AFG IRN TUR 36 CHI 37 MON 38 NKO 40 IND PAK 70 ask cdn 71 cdn 72 cdn 73 usa 74 usa 75 usa 76 usa 77 usa 78 usa 82 ven
Remarks: squarrosa agg. Lappula consanguinea (C- and E-Asian) is often kept separate
Synonyms: hidden
= Cynoglossospermum consanguineum (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Cynoglossospermum lappula (L.) Kuntze; Revis. Gen. Pl. 2: 436 (1891)
= Cynoglossum clusii Loisel.; Cuvier, Dict. Sci. Nat. ed. 2. 12: 384 (1819), nom. inval.
= Cynoglossum lappula (L.) Scop.; Fl. Carn. ed. II. 1: 125 (1771)
= Cynoglossum lappula (L.) Wallr.; 77 (1822), isonym
= Cynoglossum lusitanicum Fortis; Osserv. Sopra Cherso ed Osero: 68 (1771), nom. illeg.
= Echinospermum alpinum Schur; Enum. Pl. Transsilv.: 464 (1866)
= Echinospermum canescens var. angustifolium Opiz; Bercht., Oekon.-Techn. Fl. Böhm. 2(2): 153 (1839)
= Echinospermum canescens var. longifolium Opiz; Bercht., Oekon.-Techn. Fl. Böhm. 2(2): 153 (1839)
= Echinospermum casanense Wierzb. ex Ledeb.; Fl. Ross. 3: 156 (1847)
= Echinospermum casanicum Wierzb.; Geogr. Pl. Casan.: 84 (1842)
= Echinospermum consanguineum Fisch. & C.A.Mey.; Ind. Sem. Hort. Petrop. 5: 35 (1838)
= Echinospermum costae Sennen; Diagn. Nouv.: 302 (1936)
= Echinospermum cynoglossoides E.Mey. ex A.DC.; Prodr. [A. DC.] 10: 137 (1846)
= Echinospermum fabrei Sennen; Bull. Soc. Bot. France 73: 658 (1926 publ. 1927)
= Echinospermum fremontii Torr. & A.Gray; Pacif. Railr. Rep. Stevens, Bot. 12(2): 46 (1860)
= Echinospermum ispahanicum Boiss.; Diagn. Pl. Orient. ser. 1, 11: 123 (1849)
= Echinospermum lanceolatum Opiz; Bercht., Oekon.-Techn. Fl. Böhm. 2(2): 152 (1839)
= Echinospermum lappula (L.) Lehm.; Pl. Asperif. Nucif.: 12 1. (1818)
= Echinospermum lappula subsp. squarrosum (Retz.) Schübl. & G.Martens; 126 (1834)
= Echinospermum lappula var. consanguineum (Fisch. & C.A.Mey.) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 89 (1868)
= Echinospermum lappula var. glaucescens Wimm. & Grab.; Fl. Siles. 1: 158 (1827)
= Echinospermum lappula var. longifolium Grognot; Carion, Cat. Rais. Pl. Vasc. Départ. Saône-et-Loire: 176 (1865)
= Echinospermum lappula var. pedicellatum Rouy; Rouy & Foucaud, Fl. France 10: 341 (1908)
= Echinospermum lappula var. squarrosum (Retz.) Gaudin; Fl. Helv. 2: 59 (1828)
= Echinospermum lappula var. squarrosum (Retz.) Lehm.; Pl. Asperif. Nucif.: 121 (1818)
= Echinospermum lappula var. virescens Wimm. & Grab.; Fl. Siles. 1: 158 (1827)
= Echinospermum obtusifolium Opiz; Bercht., Oekon.-Techn. Fl. Böhm. 2(2): 150 (1839)
= Echinospermum pedunculatum Opiz; Bercht., Oekon.-Techn. Fl. Böhm. 2(2): 152 (1839)
= Echinospermum squarrosum (Retz.) Bluff & Fingerh.; 234 (1825)
= Echinospermum squarrosum (Retz.) Rchb.; Fl. Germ. Excurs. 345 (1831)
= Echinospermum vulgare Sw. ex Schrad.; Comm. Götting. 4: 195, f. 7 (1816 publ. 1818)
= Lappula consanguinea (Fisch. & C.A.Mey.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
= Lappula consanguinea var. cupuliformis C.J.Wang; Bull. Bot. Res., Harbin l(4): 88 (1981)
= Lappula echinata Gilib.; Fl. Lit. Inch. 1: 25 (1782), opus utique oppr.
= Lappula echinata Delarbre; Fl. Auvergne, éd. 2: 13 (1800), nom. illeg.
= Lappula echinata Gilib. ex Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 213 (1887), nom. superfl.
= Lappula echinata var. consanguinea (Fisch. & C.A.Mey.) Brand; Engl., Pflanzenr., IV, 252(97): 140 (1931)
= Lappula echinata var. erecta (A.Nelson) Brand; Engl., Pflanzenr., IV, 252: 140 (1931)
= Lappula echinata var. grandiflora Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 214 (1887)
= Lappula echinata var. mollita Murr; 244 (1923)
= Lappula erecta A.Nelson; Bull. Torrey Bot. Club, 268 (1900)
= Lappula fremontii (Torr. & A.Gray) Greene; Pittonia 4: 96 (1899)
= Lappula lappula (L.) H.Karst.; Ill. Repert. Pharm.-Med. Bot.: 979 (1886), nom. inval.
= Lappula myosotis f. triserialis (Trautv.) Kotejowa; Pawl., 180 (1963), nom. inval.
= Lappula myosotis Moench; Methodus: 417 (1794)
= Lappula squarrosa var. erecta (A.Nelson) Dorn; Vasc. Pl. Wyoming: 295 (1988)
= Myosotis echinata L.; Sp. Pl. 131 (1753)
= Myosotis echinosperma Mazziari; Ionios Antologia 4: 960 (1835)
= Myosotis lappula L.; Sp. Pl. 1: 131 (1753)
= Myosotis squarrosa Retz.; Obs. Bot. 2: 9 (1781)
= Rochelia lappula (L.) Roem. & Schult.; Syst. 4: 109 (1819)
species :
Lappula stricta (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
Distribution: Siberia (Altai, Buryatia, Krasnoyarsk, Tuva, W-Siberia); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang, Nei Mongol); Mongolia
30 SIB 32 KAZ KGZ TZK UZB 36 CHI 37 MON
Synonyms: hidden
= Cynoglossospermum strictum (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum strictum Ledeb.; Fl. Altaic. 1: 200 (1829)
= Hackelia stricta (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
variant :
var. leiocarpa Popov; Kom., Fl. URSS 19: 440 (1953)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Gansu, Xinjiang)
32 KAZ KGZ TZK UZB 36 CHI
species :
Lappula subcaespitosa Popov ex Golosk.; Fl. Rast. Res. Kaz.: 75 (1975)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Lappula subcaespitosa (Popov) Pavlov; Sovietsk. Bot., No. 1, 27, in obs (1934)
species :
Lappula tadshikorum Popov; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 14: 319 (1951)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang)
32 KAZ KGZ TZK UZB 36 CHI
species :
Lappula tenuis (Ledeb.) Gürke; Engl. & Prantl, Nat. Pflanzenfam. 4(3a): 107 (1897)
Distribution: Siberia (Altai, Irkutsk, Krasnoyarsk, Tuva); Kazakhstan; Uzbekistan; ?Kyrgyzstan; Tajikistan; China (Xinjiang); Mongolia
30 SIB 32 KAZ TZK UZB 36 CHI 37 MON
Synonyms: hidden
= Cynoglossospermum tenue (Ledeb.) Kuntze; Revis. Gen. Pl. 2: 437 (1891)
= Echinospermum redowskii var. tenue (Ledeb.) Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 84 (1868)
= Echinospermum tenue Ledeb.; Fl. Altaic. 1: 201 (1829)
= Hackelia tenuis (Ledeb.) Opiz; Oekon.-Techn. Fl. Böhm. [Berchtold et al.] 2(2): 147, adnot. (1839)
= Myosotis tenuis (Ledeb.) Morch; Cat. Hort. Hafn.: 64 (1839)
species :
Lappula tianschanica Popov & Zakirov; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 14: 317 (1951)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang)
32 KAZ KGZ TZK UZB 36 CHI
variant :
var. altaica C.J.Wang; Bull. Bot. Res., Harbin l(4): 86 (1981)
Distribution: China (N-Xinjiang)
36 CHI
variant :
var. gracilis C.J.Wang; Bull. Bot. Res., Harbin l(4): 86 (1981)
Distribution: China (N-Xinjiang)
36 CHI
species :
Lappula transalaica (B.Fedtsch. ex Popov) Nabiev; Consp. Fl. As. Med. 8: 198 (1986)
Distribution: Kyrgyzstan; Tajikistan; China (W-Xinjiang)
32 KGZ TZK 36 CHI
Synonyms: hidden
= Lepechiniella transalaica B.Fedtsch. ex Popov; Kom., Fl. URSS 19: 714 (1953)
species :
Lappula turczaninowii Ovchinnikova; Rast. Mir Aziatsk. Rossii 16(2): 131 (2023)
Distribution: Siberia (Irkursk, Buryatia, Yakutia); Mongolia
30 SIB 37 MON
species :
Lappula tuvinica Ovczinnikova; Fl. Sibir. (Pyrolac.-Lamiac.) 11: 142 (1997)
Distribution: Siberia (Altai, Tuva, W-Siberia); Mongolia
30 SIB 37 MON
species :
Lappula umbellulifera (B.Fedtsch. ex Popov) Sennikov; Checkl. Vasc. Pl. Tian-Shan 315 (2021)
Distribution: Tajikistan (Tianshan)
32 TZK
Conservation Status: IUCN I
Synonyms: hidden
= Tianschaniella umbellulifera B.Fedtsch.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 14: 338 (1951)
species :
Lappula zaissanica (Aralbaev) Aralbaev; Novon 5(2): 119 (1995)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Lappula coronifera subsp. zaissanica Aralbaev; Bot. Zhurn. 75(5): 707 (1990)
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A complete, synonymic checklist of the Higher Plants of the World

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Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
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